Table 2 Detailed information for 188 Triticum and Aegilops germplasm collected from GRIN platform through USDA
No. | ID | Plant name | Taxon | Country | Selection criteria | Growth habit | Probable market class |
|---|---|---|---|---|---|---|---|
1 | PI 542274 | 84TK081-057 | A. speltoides var. speltoides | Turkey | Center of origin | NA | NA |
2 | PI 560529 | TU85-008-01 | A. speltoides var. speltoides | Turkey | Center of origin | NA | NA |
3 | PI 449338 | AE 081D | A. speltoides var. speltoides | Israel | Center of domestication | NA | NA |
4 | PI 560528 | TU85-015-02-2 | A. speltoides var. ligustica | Turkey | Center of origin | NA | NA |
5 | PI 603233 | TA 1669 | A. tauschii | Azerbaijan | D genome donor | NA | NA |
6 | CItr 180 | China | T. aestivum subsp. aestivum | China | Least breast cancer, highest producer of wheat | W | SRW |
7 | CItr 14319 | India Hybrid 65 | T. aestivum subsp. aestivum | India | Least breast cancer, second highest producer of wheat | S | SWS |
8 | CItr 15136 | American 378 | T. aestivum subsp. aestivum | Sudan | Northern Africa | S | SWS |
9 | PI 9871 | Erivan | T. aestivum subsp. aestivum | Armenia | Center of origin | S | HRS |
10 | PI 9872 | Galgalos | T. aestivum subsp. aestivum | Armenia | Center of origin | S | SWS |
11 | PI 52323 | Little Joss | T. aestivum subsp. aestivum | UK | Northern Europe | W | SRW |
12 | PI 54431 | Triminia | T. aestivum subsp. aestivum | Libya | Middle East | S | HRS |
13 | PI 62004 | NA | T. aestivum subsp. aestivum | China | Least breast cancer | F | NA |
14 | PI 81791 | Sapporo Haru Komugi Jugo | T. aestivum subsp. aestivum | Japan | Eastern Asia | S | SWS |
15 | PI 82469 | Poubiru | T. aestivum subsp. aestivum | North Korea | Eastern Asia | S | HRS |
16 | PI 87117 | Ejuiea | T. aestivum subsp. aestivum | South Korea | Eastern Asia | W | HRW |
17 | PI 94418 | 99 | T. aestivum subsp. aestivum | Russia | Fourth highest producer of wheat | W | HRW |
18 | PI 116232 | Solid Straw Tuscan | T. aestivum subsp. aestivum | NZ | High cancer incidence | W | SWW |
19 | PI 124818 | Cross No. 7 | T. aestivum subsp. aestivum | NZ | High cancer incidence | S | HRS |
20 | PI 139599 | Egypt NA 101 | T. aestivum subsp. aestivum | Egypt | Western Asia | S | SWS |
21 | PI 155315 | Yemen | T. aestivum subsp. aestivum | Yemen | Western Asia | S | SRS |
22 | PI 165208 | Sert Bolvadin | T. aestivum subsp. aestivum | Turkey | Center of origin | S | SWS |
23 | PI 178383 | 6256 | T. aestivum subsp. aestivum | Turkey | Center of origin | W | SRW |
24 | PI 184994 | Snogg II | T. aestivum subsp. aestivum | Norway | Northern Europe | S | HRS |
25 | PI 190451 | Snogg | T. aestivum subsp. aestivum | Norway | Northern Europe | S | SRS |
26 | PI 191334 | Marzuolo 87 | T. aestivum subsp. aestivum | Italy | Southern Europe | W | HWW |
27 | PI 265482 | Olympia | T. aestivum subsp. aestivum | Finland | Northern Europe | S | SWS |
28 | PI 266879 | S995 | T. aestivum subsp. aestivum | Iraq | Center of origin | S | HWS |
29 | PI 266880 | S997 | T. aestivum subsp. aestivum | Iraq | Center of origin | S | HRS |
30 | PI 274505 | NA | T. aestivum subsp. aestivum | Thailand | South Eastern Asia | S | SWS |
31 | PI 283150 | Horani Nawawi | T. aestivum subsp. aestivum | Jordan | Center of origin | S | SWS |
32 | PI 297005 | AFRICA MAYO | T. aestivum subsp. aestivum | Kenya | Eastern Africa | S | HRS |
33 | PI 347003 | White Shanazi | T. aestivum subsp. aestivum | Afghanistan | South Central Asia | S | SWS |
34 | PI 350308 | DACIA | T. aestivum subsp. aestivum | Romania | Eastern Asia | W | HRW |
35 | PI 384399 | Nigeria-2 | T. aestivum subsp. aestivum | Nigeria | Western Africa | S | HWS |
36 | PI 406475 | 2 | T. aestivum subsp. aestivum | Nepal | South Central Asia | S | HRS |
37 | PI 414975 | No. 6 | T. aestivum subsp. aestivum | Indonesia | South Central Asia | S | HRS |
38 | PI 480034 | MG 31147 | T. aestivum subsp. aestivum | Ethiopia | Eastern Africa | S | SWS |
39 | PI 487292 | SY 270 | T. aestivum subsp. aestivum | Jordan | Center of origin | S | HRS |
40 | PI 519554 | PAKISTAN 20 | T. aestivum subsp. aestivum | Kenya | Eastern Africa | S | SWS |
41 | PI 532053 | 96 | T. aestivum subsp. aestivum | Egypt | Northern Africa | S | SWS |
42 | PI 585019 | 15007 | T. aestivum subsp. aestivum | Saudi Arabia | Western Asia | S | HRS |
43 | PI 585024 | 15063 | T. aestivum subsp. aestivum | Saudi Arabia | Western Asia (a part of Core 95) | S | HRS |
44 | PI 603919 | UCRBW98-2 | T. aestivum subsp. aestivum | USA | DNA segment from Spelta and HWS Pavon (bread wheat), Northern America, a part of core 6 | S | SWS |
45 | PI 648392 | KUNDAN | T. aestivum subsp. aestivum | India | Least breast cancer, second highest producer of wheat | S | HWS |
46 | CItr 14352 | II-50-25 | T. aestivum subsp. aestivum | Paraguay | South America | S | HRS |
47 | PI 10611 | Talimka | T. aestivum subsp. aestivum | Turkmenistan | South Central Asia | S | HWS |
48 | PI 61693 | 294 | T. aestivum subsp. aestivum | Malawi | Eastern Africa | S | HWS |
49 | PI 91235 | Cagayan | T. aestivum subsp. aestivum | Philippines | South East Asia | S | HRS |
50 | PI 125088 | Ile de France | T. aestivum subsp. aestivum | France | Western Europe, more cancer | S | HWS |
51 | PI 174657 | Ile de France | T. aestivum subsp. aestivum | France | Western Europe, more cancer | S | SRS |
52 | PI 182665 | 9915 | T. aestivum subsp. aestivum | Lebanon | Western Asia | S | HWS |
53 | PI 191701 | B 256 F.S. 1354 | T. aestivum subsp. aestivum | Mozambique | Eastern Africa | F | SWS |
54 | PI 191744 | FL S Aurora | T. aestivum subsp. aestivum | Mozambique | Eastern Africa | S | SWS |
55 | PI 203081 | Sabanero | T. aestivum subsp. aestivum | Tanzania | Eastern Africa | S | HRS |
56 | PI 231115 | II-2734-2c(1–2)x2T | T. aestivum subsp. aestivum | Guatemala | Central America | S | HRS |
57 | PI 234233 | Idaho 1877 NR AE | T. aestivum subsp. aestivum | Zambia | Eastern Africa | S | HRS |
58 | PI 278386 | Morocco 58 | T. aestivum subsp. aestivum | Morocco | Northern Africa | S | HRS |
59 | PI 278395 | Poland 2 | T. aestivum subsp. aestivum | Poland | Eastern Europe | W | HRW |
60 | PI 313098 | Quern | T. aestivum subsp. aestivum | Ireland | Northern Europe | S | HRS |
61 | PI 344018 | Mistura Pinto | T. aestivum subsp. aestivum | Angola | Middle Africa, least cancers | S | SWS |
62 | PI 344019 | Saraiva Vieira | T. aestivum subsp. aestivum | Angola | Middle Africa, least cancers | S | SWS |
63 | PI 351474 | Reval | T. aestivum subsp. aestivum | Estonia | Northern Europe | W | HRW |
64 | PI 351870 | 10180-54-29 | T. aestivum subsp. aestivum | Burundi | Eastern Africa | S | SWS |
65 | PI 374248 | BOL-17 | T. aestivum subsp. aestivum | Chad | Middle Africa, least cancers | S | HRS |
66 | PI 374249 | BOL-19 | T. aestivum subsp. aestivum | Chad | Middle Africa, least cancers | S | HRS |
67 | PI 374254 | 34335 | T. aestivum subsp. aestivum | Mali | Western Africa | S | HWS |
68 | PI 384399 | Nigeria-2 | T. aestivum subsp. aestivum | Nigeria | Western Africa | S | HWS |
69 | PI 410425 | ILICHEVKA | T. aestivum subsp. aestivum | Kazakhstan | South Central Asia | W | SRW |
70 | PI 428690 | LEUCURUM 3 | T. aestivum subsp. aestivum | Uzbekistan | South Central Asia | S | SRS |
71 | PI 470905 | MG 18060 | T. aestivum subsp. aestivum | Algeria | Northern Africa | S | SWS |
72 | PI 480481 | R-124 | T. aestivum subsp. aestivum | Bolivia | South America | S | SRS |
73 | PI 481713 | 41 | T. aestivum subsp. aestivum | Bhutan | South Central Asia | W | HRW |
74 | PI 481715 | Ka | T. aestivum subsp. aestivum | Bhutan | South Central Asia | S | SWS |
75 | PI 486155 | CHIWORE | T. aestivum subsp. aestivum | Zimbabwe | Eastern Africa, least breast cancer | S | HRS |
76 | PI 486156 | GWEBI | T. aestivum subsp. aestivum | Zimbabwe | Eastern Africa, least breast cancer | S | SWS |
77 | PI 486157 | RUSAPE | T. aestivum subsp. aestivum | Zimbabwe | Eastern Africa, least breast cancer | S | HRS |
78 | PI 490405 | Koira alkuna | T. aestivum subsp. aestivum | Mali | Western Africa | S | HWS |
79 | PI 494926 | ZFA 3145 | T. aestivum subsp. aestivum | Zambia | Eastern Africa | S | HRS |
80 | PI 532301 | Alas | T. aestivum subsp. aestivum | Oman | Western Asia | S | SRS |
81 | PI 573754 | NSGC 531 | T. aestivum subsp. aestivum | Hondurus | Central America | S | HRS |
82 | PI 591964 | EMBRAPA 16 | T. aestivum subsp. aestivum | Brazil | South America | S | HRS |
83 | PI 639354 | TJK03-128 | T. aestivum subsp. aestivum | Tazikistan | South Central Asia | S | SWS |
84 | PI 648894 | Dickson's No. 444 | T. aestivum subsp. aestivum | Argentina | South America | S | HWS |
85 | Cltr 14108 | Chinese Spring | T. aestivum subsp. aestivum | USA | Northern America | S | SRS |
86 | PI 434642 | TINCURRIN | T. aestivum subsp. compactum | Australia | More cancer especially skin cancer | S | Club wheat |
87 | PI 190963 | Spelta Hohenheim | T. aestivum subsp. spelta | Portugal | Southern Europe | S | Spelt |
88 | PI 348710 | 69Z6.894 | T. aestivum subsp. spelta | Spain | Southern Europe | S | Spelt |
89 | PI 355625 | Spelta 34 | T. aestivum subsp. spelta | Belgium | Western Europe | S | Spelt |
90 | PI 591895 | NA | T. aestivum subsp. spelta | Germany | Central Europe | W | Spelt |
91 | PI 367199 | 128 | T. aestivum subsp. spelta | Afghanistan | Central Asia | W | Spelt |
92 | PI 538510 | G2830 | T. monococcum subsp. aegilopoides | Iraq | Center of origin, Western Asia | S | Wild einkorn |
93 | PI 167526 | 2485 | T. monococcum subsp. aegilopoides | Turkey | Western Asia | S | Wild einkorn |
94 | PI 266844 | 87 | T. monococcum subsp. aegilopoides | Uk | Northern Europe | S | Wild einkorn |
95 | PI 427990 | G3114 | T. monococcum subsp. aegilopoides | Lebanon | Center of domestication | W | Wild einkorn |
96 | PI 427304 | G1764 | T. timopheevii subsp. armeniacum | Armenia | Center of origin, Western Asia | W | Wild timopheevii |
97 | PI 538478 | G2633 | T. timopheevii subsp. armeniacum | Iraq | Center of domestication | W | Wild timopheevii |
98 | PI 70738 | 22 | T. turgidum subsp. carthlicum | Iraq | Center of origin, Western Asia | S | Persian wheat |
99 | PI 532501 | H83-1537 | T. turgidum subsp. carthlicum | Soviet Union | Â | S | Persian wheat |
100 | PI 471808 | G-485-5 M | T. turgidum subsp. dicoccoides | Israel | Center of domestication | W | Wild emmer |
101 | PI 471778 | G-40-1-2B-1 M | T. turgidum subsp. dicoccoides | Israel | Center of domestication | W | Wild emmer |
102 | PI 2789 | Yaroslav Spring | T. turgidum subsp. dicoccon | Russia | Fourth highest producer of wheat | S | Cultivated emmer |
103 | PI 73388 | 2868 | T. turgidum subsp. dicoccon | Armenia | Center of origin, Western Asia | S | Cultivated emmer |
104 | PI 79899 | N-64 | T. turgidum subsp. dicoccon | China | Least breast cancer, highest producer of wheat | S | Cultivated emmer |
105 | PI 190920 | 2323A | T. turgidum subsp. dicoccon | Portugal | Southern Europe | S | Cultivated emmer |
106 | PI 191390 | Rufum | T. turgidum subsp. dicoccon | Ethiopia | Eastern Africa | S | Cultivated emmer |
107 | PI 308879 | NA | T. turgidum subsp. dicoccon | Spain | Southern Europe | S | Cultivated emmer |
108 | PI 355483 | T 563 | T. turgidum subsp. dicoccon | Spain | Southern Europe | S | Cultivated emmer |
109 | PI 355485 | T 567 | T. turgidum subsp. dicoccon | Spain | Southern Europe | S | Cultivated emmer |
110 | PI 499973 | KU 1533 | T. turgidum subsp. dicoccon | Armenia | Center of origin, Western Asia | S | Cultivated emmer |
111 | PI 154582 | NA | T. turgidum subsp. dicoccon | Taiwan | Eastern Asia | S | Cultivated emmer |
112 | CItr 15185 | Hudeiba 154 | T. turgidum subsp. durum | Sudan | Northern Africa | S | Durum or macaroni |
113 | PI 9130 | Saragolla | T. turgidum subsp. durum | Italy | Southern Europe | S | Durum or macaroni |
114 | PI 54432 | Tripshiro | T. turgidum subsp. durum | Libya | Northern Africa | S | Durum or macaroni |
115 | PI 67341 | Huguenot | T. turgidum subsp. durum | Australia | More cancer especially skin cancer | S | Durum or macaroni |
116 | PI 78809 | CI 10107 | T. turgidum subsp. durum | Georgia | Western Asia | S | Durum or macaroni |
117 | PI 81792 | Marching No. 8 | T. turgidum subsp. durum | Japan | Eastern Asia | S | Durum or macaroni |
118 | PI 94701 | 390 | T. turgidum subsp. durum | Ancient Palestine | Western Asia | S | Durum or macaroni |
119 | PI 133459 | Durum H2 | T. turgidum subsp. durum | Egypt | Center of origin, Northern Africa | S | Durum or macaroni |
120 | PI 153726 | Sicilian | T. turgidum subsp. durum | N. Africa | Less cancers | S | Durum or macaroni |
121 | PI 157955 | Francesone | T. turgidum subsp. durum | Italy | Southern Europe | S | Durum or macaroni |
122 | PI 174645 | Huguenot | T. turgidum subsp. durum | Australia | More cancer especially skin cancer | S | Durum or macaroni |
123 | PI 182113 | S-44 | T. turgidum subsp. durum | Pakistan | South Central Asia | S | Durum or macaroni |
124 | PI 184532 | Russia | T. turgidum subsp. durum | Russia | Europe | S | Durum or macaroni |
125 | PI 208908 | Mendola | T. turgidum subsp. durum | Iraq | Center of origin | S | Durum or macaroni |
126 | PI 208910 | Sin El-Jamil | T. turgidum subsp. durum | Iraq | Center of origin | S | Durum or macaroni |
127 | PI 210848 | 7979 | T. turgidum subsp. durum | Iran | South Central Asia | S | Durum or macaroni |
128 | PI 221702 | 11 | T. turgidum subsp. durum | Indonesia | Southern Eastern Asia | S | Durum or macaroni |
129 | PI 231380 | Saragolla | T. turgidum subsp. durum | Italy | Oldest durum variety in Italy | S | Durum or macaroni |
130 | PI 261823 | Namra | T. turgidum subsp. durum | Saudi Arabia | Western Asia | S | Durum or macaroni |
131 | PI 265017 | 796 | T. turgidum subsp. durum | Serbia | Southern Europe | S | Durum or macaroni |
132 | PI 278223 | Gartons Early Cone | T. turgidum subsp. durum | UK | Northern Europe | W | Durum or macaroni |
133 | PI 278258 | Greece 1 | T. turgidum subsp. durum | Greece | Southern Europe | S | Durum or macaroni |
134 | PI 278509 | Valencia 6 | T. turgidum subsp. durum | Spain | Southern Europe | S | Durum or macaroni |
135 | PI 278553 | Tripolitco | T. turgidum subsp. durum | Cyprus | Western Asia | S | Durum or macaroni |
136 | PI 283853 | China 34 | T. turgidum subsp. durum | China | Least breast cancer, highest producer of wheat | S | Durum or macaroni |
137 | PI 306571 | R.S.N. | T. turgidum subsp. durum | Italy | Southern Europe | S | Durum or macaroni |
138 | PI 325850 | PW 3 | T. turgidum subsp. durum | India | Least breast cancer, second highest producer of wheat | S | Durum or macaroni |
139 | PI 361149 | Bijaga Yellow | T. turgidum subsp. durum | India | Least breast cancer, second highest producer of wheat | S | Durum or macaroni |
140 | PI 362046 | C 1138/63 | T. turgidum subsp. durum | Romania | Eastern Europe | W | Durum or macaroni |
141 | PI 422295 | QUILAFEN | T. turgidum subsp. durum | Chile | South America | S | Durum or macaroni |
142 | PI 422297 | SINCAPE 90 | T. turgidum subsp. durum | Italy | Southern Europe | S | Durum or macaroni |
143 | PI 422312 | MACS-45 | T. turgidum subsp. durum | India | Least breast cancer, second highest producer of wheat | S | Durum or macaroni |
144 | PI 428458 | Egypt Local No. 8 | T. turgidum subsp. durum | Egypt | Center of origin, Northern Africa | S | Durum or macaroni |
145 | PI 428468 | JORDAN 38 | T. turgidum subsp. durum | Jordan | Center of origin, Western Asia | S | Durum or macaroni |
146 | PI 428469 | JORDAN 40 | T. turgidum subsp. durum | Jordan | Center of origin, Western Asia | S | Durum or macaroni |
147 | PI 462107 | 172 | T. turgidum subsp. durum | Yemen | Western Asia | S | Durum or macaroni |
148 | PI 480347 | MG 31577 | T. turgidum subsp. durum | Ethiopia | Eastern Africa | S | Durum or macaroni |
149 | PI 496260 | MEDORA | T. turgidum subsp. durum | Canada | North America | S | Durum or macaroni |
150 | PI 519864 | DURUM VARIETY 24 | T. turgidum subsp. durum | Mexico | North America | S | Durum or macaroni |
151 | PI 520393 | TUNISIAN DURUM 1 | T. turgidum subsp. durum | Tunisia | Northern Africa | S | Durum or macaroni |
152 | PI 520394 | TUNISIAN DURUM 8 | T. turgidum subsp. durum | Tunisia | Northern Africa | S | Durum or macaroni |
153 | PI 520414 | ICD 7780-5AP-OSH-OAP | T. turgidum subsp. durum | Syria | Center of origin, Western Asia | S | Durum or macaroni |
154 | PI 520415 | SYRIAN DURUM 27 | T. turgidum subsp. durum | Syria | Center of origin, Western Asia | S | Durum or macaroni |
155 | PI 542464 | SHORT SARAGOLLA | T. turgidum subsp. durum | USA | High cancer incidence | S | Durum or macaroni |
156 | PI 585025 | 15017 | T. turgidum subsp. durum | Saudi Arabia | Western Asia | S | Durum or macaroni |
157 | CItr 14374 | 497-360 | T. turgidum subsp. durum | Lebanon | Western Asia | S | Durum or macaroni |
158 | CItr 14802 | ELS 6404-122 | T. turgidum subsp. durum | Eritrea | Eastern Africa | S | Durum or macaroni |
159 | PI 5465 | Candeal | T. turgidum subsp. durum | Argentina | South America | F | Durum or macaroni |
160 | PI 5639 | Kubanka | T. turgidum subsp. durum | Kazakhstan | South Central Asia, largest consumer of wheat | S | Durum or macaroni |
161 | PI 35314 | 1809a | T. turgidum subsp. durum | Kyrgyzstan | South Central Asia | F | Durum or macaroni |
162 | PI 50929 | 933 | T. turgidum subsp. durum | Kyrgyzstan | South Central Asia | S | Durum or macaroni |
163 | PI 61108 | 6951 | T. turgidum subsp. durum | Turkmenistan | South Central Asia | S | Durum or macaroni |
164 | PI 89642 | NA | T. turgidum subsp. durum | Hondurus | Central America | S | Durum or macaroni |
165 | PI 278384 | Morocco C10895 | T. turgidum subsp. durum | Morocco | Northern Africa | W | Durum or macaroni |
166 | PI 286066 | NA | T. turgidum subsp. durum | Poland | Eastern Europe | S | Durum or macaroni |
167 | PI 384401 | Wurno 2 | T. turgidum subsp. durum | Nigeria | Western Africa | S | Durum or macaroni |
168 | PI 519759 | D 73121 | T. turgidum subsp. durum | Algeria | Northern Africa | S | Durum or macaroni |
169 | PI 520164 | ALGERIA LINE 47 | T. turgidum subsp. durum | Algeria | Northern Africa | S | Durum or macaroni |
170 | PI 532289 | Musane | T. turgidum subsp. durum | Oman | Western Asia | S | Durum or macaroni |
171 | PI 565208 | Chaggo | T. turgidum subsp. durum | Bolivia | South America | S | Durum or macaroni |
172 | PI 592019 | VATAN | T. turgidum subsp. durum | Uzbekistan | South Central Asia | S | Durum or macaroni |
173 | PI 654290 | TJK2006:296 | T. turgidum subsp. durum | Tazikistan | South Central Asia | S | Durum or macaroni |
174 | Cltr 13165 | Langdon | T. turgidum subsp. durum | USA | Northern America | W | Durum or macaroni |
175 | PI 330553 | 189 | T. turgidum subsp. paleocolchicum | UK | Northern Europe | S | Cultivated emmer |
176 | PI 211708 | Egypt | T. turgidum subsp. turanicum | Egypt | Center of origin, Northern Africa | S | Khorasan or oriental |
177 | PI 166591 | Ak | T. turgidum subsp. turgidum | Turkey | Center of origin | S | Rivet or cone |
178 | PI 167867 | 4314 | T. turgidum subsp. turgidum | Turkey | Center of origin | S | Rivet or cone |
179 | PI 481591 | IQ 223 | T. turgidum subsp. turgidum | Iraq | Center of origin | S | Rivet or cone |
180 | PI 502933 | Fo Shou Mai | T. turgidum subsp. turgidum | China | Largest producer | S | Rivet or cone |
181 | PI 208912 | Zerdakia | T. turgidum subsp. turgidum | Iraq | Center of origin | S | Rivet or cone |
182 | PI 438971 | AKMOLINKA 2 | T. turgidum subsp. turgidum | Kazakhstan | South Central Asia | S | Rivet or cone |
183 | PI 427328 | G2264 | T. urartu | Iraq | Center of origin | S | Wild einkorn |
184 | PI 428183 | G1759 | T. urartu | Armenia | Center of origin | S | Wild einkorn |
185 | PI 428279 | G3162 | T. urartu | Lebanon | Center of domestication | W | Wild einkorn |
186 | PI 355707 | 69Z5.72 | T. zhukovskyi | Georgia | Donor for GG genome and cross between T. timopheevi and T. monococcum | W | Cultivated hexaploid |
187 | PI 429099 | 6A-696 | Triticosecale sp. | Germany | Cross between T. dicoccum & S. cereale; hexaploid | F | Tritical (Rye and durum cross) |
188 | PI 574284 | ASVM4*4654 | T. hybrid | USA | High cancer incidence | S | Aegilops squarrosa/T. dicoccum |